主要从事早期动物起源和寒武纪大爆发时期的古生态学、古生物学、地层学及其环境背景的研究。工作重点集中在中国寒武纪特异埋藏化石库（如中国澄江生物群、关山生物群等）的古生态学、埋藏学和系统古生物学研究。

   

070903-古生物学与地层学

寒武纪特异埋藏化石群
古生态学
寒武纪大爆发与生命演化

2006-09--2009-07   中国科学院南京地质古生物研究所   博士学位
2003-09--2006-07   中国科学院南京地质古生物研究所   硕士学位
1999-09--2003-07   中国地质大学（北京）   学士学位

研究生学历

博士学位

   

2017-03~现在, 中国科学院南京地质古生物研究所, 研究员
2013-01~2017-03,中国科学院南京地质古生物研究所, 副研究员
2012-07~2013-07,美国南加州大学(University of Southern California), 访问学者
2010-12~2011-05,加拿大安大略皇家自然博物馆(Royal Ontario Museum), 访问学者
2009-08~2012-12,中国科学院南京地质古生物研究所, 助理研究员

   

（1） 中国地质学会2016年度十大地质科技进展, , 其他, 2016
（2） 中国科学院青年创新促进会会员, , 院级, 2013
（3） 杰出青年工作者, , 研究所（学校）, 2013

   

[1] 孙智新, 杨爱华, 赵方臣. New trilobite assemblage from the lower Cambrian (upper Stage 4) of the Lake Zone, western Mongolia. Journal of Paleontology[J]. 2023, [2] Sun, Zhixin, Zhao, Fangchen, Zeng, Han, Luo, Cui, Van Iten, Heyo, Zhu, Maoyan. The middle Cambrian Linyi Lagerstatte from the North China Craton: a new window on Cambrian evolutionary fauna. NATIONAL SCIENCE REVIEW[J]. 2022, 9(7): 146-155, [3] Wang, DeZhi, Zhao, FangChen, Zeng, Han, Li, GuoXiang, Zhu, MaoYan. A new rhynchonelliform brachiopod Longtancunella with soft-part preservation from the Hongjingshao Formation (Cambrian Stage 3) in Yunnan, South China. PALAEOWORLD[J]. 2022, 31(4): 570-581, http://dx.doi.org/10.1016/j.palwor.2021.12.002.
[4] Zeng, H., 赵方臣, Zhu, M.Y.. Innovatiocaris, a complete radiodont from the early Cambrian Chengjiang Lagerstatte and its implications for the phylogeny of Radiodonta. JOURNAL OF THE GEOLOGICAL SOCIETY[J]. 2022, [5] Sun, Zhixin, Zeng, Han, Zhao, Fangchen. Unique Cambrian trilobite Eodontopleura: an example of strong convergence in trilobite evolution. HISTORICAL BIOLOGY. 2022, [6] Tian, Qingyi, Zhao, Fangchen, Zeng, Han, Zhu, Maoyan, Jiang, Baoyu. Ultrastructure reveals ancestral vertebrate pharyngeal skeleton in yunnanozoans. SCIENCE[J]. 2022, 377(6602): 218-+, [7] Sun, H.J., 赵方臣, Zhu, M.Y.. Anatomy, Paleoautecology, and Phylogentic affinity of tubular Glossolites magnus from the early Cambrian Chengjiang Biota, South China. Paper in Palaeontology[J]. 2022, [8] Aria, Cedric, Zhao, Fangchen, Zhu, Maoyan. Fuxianhuiids are mandibulates and share affinities with total-group Myriapoda. JOURNAL OF THE GEOLOGICAL SOCIETY[J]. 2021, 178(5): http://dx.doi.org/10.1144/jgs2020-246.
[9] 陈湘远, 赵方臣, 杨爱华. 埃迪卡拉纪至寒武纪苗岭世刺细胞动物化石研究现状与展望. 古生物学报[J]. 2021, 60(1): 87-107, http://lib.cqvip.com/Qikan/Article/Detail?id=7104556437.
[10] 刘瑶, 孙智新, 曾晗, 赵方臣. 山东潍坊寒武系苗岭统乌溜阶馒头组上部的腕足动物. 古生物学报[J]. 2021, 60(1): 150-165, http://lib.cqvip.com/Qikan/Article/Detail?id=7104556441.
[11] 殷宗军, 赵方臣. 动物起源和寒武纪大爆发——序言. 古生物学报[J]. 2021, 60(1): 1-9, http://lib.cqvip.com/Qikan/Article/Detail?id=7104556432.
[12] 孙智新, 曾晗, 赵方臣. 山东寒武系苗岭统三叶虫消化系统的化石证据. 古生物学报[J]. 2021, 60(1): 166-175, https://kns.cnki.net/KCMS/detail/detail.aspx?dbcode=CJFQ&dbname=CJFDLAST2021&filename=GSWX202101011&v=MTA4OTRxVHJXTTFGckNVUjd1ZlllZG1GQ3JnVTdyTklqN2Nkckc0SE5ETXJvOUVaWVI4ZVgxTHV4WVM3RGgxVDM=.
[13] Zeng, Han, Zhao, FangChen, Yin, ZongJun, Zhu, MaoYan. A new early Cambrian bivalved euarthropod from Yunnan, China and general interspecific morphological and size variations in Cambrian hymenocarines. PALAEOWORLD[J]. 2021, 30(3): 387-397, http://dx.doi.org/10.1016/j.palwor.2020.09.002.
[14] Lan, Tian, Zhao, Yuanlong, Zhao, Fangchen, He, You, Martinez, Pedro, Strausfeld, Nicholas J. Leanchoiliidae reveals the ancestral organization of the stem euarthropod brain. CURRENT BIOLOGY[J]. 2021, 31(19): 4397-+, http://dx.doi.org/10.1016/j.cub.2021.07.048.
[15] Sun, Haijing, Zhao, Fangchen, Steiner, Michael, Li, Guoxiang, Na, Lin, Pan, Bing, Yin, Zongjun, Zeng, Han, Van Iten, Heyo, Zhu, Maoyan. Skeletal faunas of the lower Cambrian Yu'anshan Formation, eastern Yunnan, China: Metazoan diversity and community structure during the Cambrian Age 3. PALAEOGEOGRAPHY PALAEOCLIMATOLOGY PALAEOECOLOGY[J]. 2020, 542: http://dx.doi.org/10.1016/j.palaeo.2019.109580.
[16] Sun, Zhixin, Zeng, Han, Zhao, Fangchen. First occurrence of the Cambrian arthropod Sidneyia Walcott, 1911 outside of Laurentia. GEOLOGICAL MAGAZINE[J]. 2020, 157(3): 405-410, https://www.webofscience.com/wos/woscc/full-record/WOS:000524942400004.
[17] 赵方臣. 寒武纪大爆发与澄江生物群. 2020, 99-, [[["https://nxgp.cnki.net/kcms/detail?v=3uoqIhG8C467SBiOvrai6ZrZfyGnw2H_RdsjEQmEf5UCbQTaMZPFdeyCKUWNKPjauv8owPoLcVCG05G9Kl_qjWFpmlhjAOPq&uniplatform=NZKPT"]]].
[18] Sun, Zhixin, Zeng, Han, Zhao, Fangchen. Occurrence of the hurdiid radiodont Cambroraster in the middle Cambrian (Wuliuan) Mantou Formation of North China. JOURNAL OF PALEONTOLOGY[J]. 2020, 94(5): 881-886, https://www.webofscience.com/wos/woscc/full-record/WOS:000561371500006.
[19] Cdric Aria, Fangchen Zhao, Han Zeng, Jin Guo, Maoyan Zhu. Fossils from South China redefine the ancestral euarthropod body plan. BMC EVOLUTIONARY BIOLOGY[J]. 2020, 20(1): 1-17, http://dx.doi.org/10.1186/s12862-019-1560-7.
[20] Bottjer, David J, Yin, Zongjun, Zhao, Fangchen, Zhu, Maoyan. Comparative taphonomy and phylogenetic signal of phosphatized Weng'an and Kuanchuanpu Biotas. PRECAMBRIAN RESEARCH[J]. 2020, 349: http://dx.doi.org/10.1016/j.precamres.2019.105408.
[21] Sun, Haijing, Yin, Zongjun, Li, Guoxiang, Zhao, Fangchen, Zeng, Han, Zhu, Maoyan. Periodic shell decollation as an ecology-driven strategy in the early Cambrian Cupitheca. PALAEONTOLOGY[J]. 2020, 63(3): 431-442, https://www.webofscience.com/wos/woscc/full-record/WOS:000501465400001.
[22] Luo, Cui, Zhao, Fangchen, Zeng, Han. The first report of a vauxiid sponge from the Cambrian Chengjiang Biota. JOURNAL OF PALEONTOLOGY[J]. 2020, 94(1): 28-33, https://www.webofscience.com/wos/woscc/full-record/WOS:000503820500003.
[23] Zeng, Han, Zhao, Fangchen, Niu, Kecheng, Zhu, Maoyan, Huang, Diying. An early Cambrian euarthropod with radiodont-like raptorial appendages. NATURE[J]. 2020, 588(7836): 101-+, https://www.webofscience.com/wos/woscc/full-record/WOS:000585787700002.
[24] 朱茂炎, 杨爱华, 袁金良, 李国祥, 张俊明, 赵方臣, SooYeun, AHN, 苗兰云. 中国寒武纪综合地层和时间框架. 中国科学:地球科学[J]. 2019, 26-65, https://www.sciengine.com/doi/10.1360/N072017-00370.
[25] Maoyan ZHU, Aihua YANG, Jinliang YUAN, Guoxiang LI, Junming ZHANG, 赵方臣, SooYeun AHN, Lanyun MIAO. Cambrian integrative stratigraphy and timescale of China. 中国科学:地球科学英文版[J]. 2019, 62(1): 25-60, http://lib.cqvip.com/Qikan/Article/Detail?id=6100172027.
[26] Zhu Maoyan, Yang Aihua, Yuan Jinliang, Li Guoxiang, Zhang Junming, Zhao Fangchen, Ahn Sooyeun, Miao Lanyun. Cambrian integrative stratigraphy and timescale of China. SCIENCE CHINA. EARTH SCIENCES[J]. 2019, 62(1): 25-60, https://www.sciengine.com/doi/10.1007/s11430-017-9291-0.
[27] 朱茂炎, 赵方臣, 殷宗军, 曾晗, 李国祥. 中国的寒武纪大爆发研究：进展与展望. 中国科学:地球科学[J]. 2019, 49(10): 1455-1490, http://lib.cqvip.com/Qikan/Article/Detail?id=7100265207.
[28] Yin, Zongjun, Zhao, Duoduo, Pan, Bing, Zhao, Fangchen, Zeng, Han, Li, Guoxiang, Bottjer, David J, Zhu, Maoyan. Early Cambrian animal diapause embryos revealed by X-ray tomography. GEOLOGY[J]. 2018, 46(5): 387-390, http://ir.nigpas.ac.cn/handle/332004/14275.
[29] 赵方臣. 东平寒武系馒头组顶部胸针球接子（Peronopsis Hawle and Corda，1847）的新材料. 古生物学报. 2018, [30] 赵方臣. Morphology of diverse radiodontan head sclerites from the early Cambrian Chengjiang Lagerstätte, southwest China. Journal of Systematic Palaeontology. 2018, [31] Ceng Han, Zhao Fangchen, Yin Zongjun, Zhu Maoyan. A new radiodontan oral cone with a unique combination of anatomical features from the early Cambrian Guanshan Lagerstätte, eastern Yunnan, South China. JOURNALOFPALEONTOLOGY[J]. 2018, 92(1): 40-48, http://www.irgrid.ac.cn/handle/1471x/1756955.
[32] Zeng, Han, Zhao, Fangchen, Yin, Zongjun, Zhu, Maoyan. Morphology of diverse radiodontan head sclerites from the early Cambrian Chengjiang Lagerstatte, south-west China. JOURNAL OF SYSTEMATIC PALAEONTOLOGY[J]. 2018, 16(1): 1-37, http://ir.nigpas.ac.cn/handle/332004/14367.
[33] 胡春林, 赵方臣, 纪秋梅, 朱茂炎. 宁镇山脉的盖帽碳酸盐岩. 地层学杂志[J]. 2018, 42(4): 381-392, http://lib.cqvip.com/Qikan/Article/Detail?id=676826819.
[34] Zeng, Han, Zhao, Fangchen, Yin, Zongjun, Zhu, Maoyan. A new radiodontan oral cone with a unique combination of anatomical features from the early Cambrian Guanshan Lagerstatte, eastern Yunnan, South China. JOURNAL OF PALEONTOLOGY[J]. 2018, 92(1): 40-48, http://ir.nigpas.ac.cn/handle/332004/14356.
[35] 董存治, 胡世学, 赵方臣. 葫芦虫化石在寒武纪关山生物群中的发现及其意义. 古生物学报[J]. 2018, 57(4): 424-428, http://lib.cqvip.com/Qikan/Article/Detail?id=6100177793.
[36] Sun, HaiJing, Zhao, FangChen, Wen, RongQin, Zeng, Han, Peng, Jin. Feeding strategy and locomotion of Cambrian hyolithides. PALAEOWORLD[J]. 2018, 27(3): 334-342, http://dx.doi.org/10.1016/j.palwor.2018.03.003.
[37] 孙智新, 王平丽, 赵方臣, 袁金良. 山东东平寒武系馒头组顶部胸针球接子（PeronopsisHawleandCorda，1847）的新材料. 古生物学报[J]. 2018, 57(3): 287-294, http://lib.cqvip.com/Qikan/Article/Detail?id=676425600.
[38] Hu, Shixue, Erdtmann, BerndD, Steiner, Michael, Zhang, Yuandong, Zhao, Fangchen, Zhang, Zhiliang, Han, Jian. Malongitubus: a possible pterobranch hemichordate from the early Cambrian of South China. JOURNAL OF PALEONTOLOGY[J]. 2018, 92(1): 26-32, http://ir.nigpas.ac.cn/handle/332004/14355.
[39] Sun, Haijing, Smith, Martin R, Zeng, Han, Zhao, Fangchen, Li, Guoxiang, Zhu, Maoyan. Hyoliths with pedicles illuminate the origin of the brachiopod body plan. PROCEEDINGS OF THE ROYAL SOCIETY B-BIOLOGICAL SCIENCES[J]. 2018, 285(1887): http://ir.nigpas.ac.cn/handle/332004/14197.
[40] Sun, Haijing, Smith, Martin R, Zeng, Han, Zhao, Fangchen, Li, Guoxiang, Zhu, Maoyan. Hyoliths with pedicles illuminate the origin of the brachiopod body plan. PROCEEDINGS OF THE ROYAL SOCIETY B-BIOLOGICAL SCIENCES[J]. 2018, 285(1887): http://ir.nigpas.ac.cn/handle/332004/14197.
[41] Wood, R A, Zhuravlev, A Yu, Sukhov, S S, Zhu, M, Zhao, F. Demise of Ediacaran dolomitic seas marks widespread biomineralization on the Siberian Platform. GEOLOGY[J]. 2017, 45(1): 27-30, http://dx.doi.org/10.1130/G38367.1.
[42] Zhu, M, Zhuravlev, A Yu, Wood, R A, Zhao, F, Sukhov, S S. A deep root for the Cambrian explosion: Implications of new bio-and chemostratigraphy from the Siberian Platform. GEOLOGY[J]. 2017, 45(5): 459-462, http://dx.doi.org/10.1130/G38865.1.
[43] 朱茂炎. A crown group priapulid from the early Cambrian Guanshan Lagerstätte. Geological Magazine. 2017, [44] Zeng, Han, Zhao, Fangchen, Yin, Zongjun, Zhu, Maoyan. Appendages of an early Cambrian metadoxidid trilobite from Yunnan, SW China support mandibulate affinities of trilobites and artiopods. GEOLOGICAL MAGAZINE[J]. 2017, 154(6): 1306-1328, [45] Zhao, Fangchen, Smith, Martin R, Yin, Zongjun, Zeng, Han, Li, Guoxiang, Zhu, Maoyan. Orthrozanclus elongata n. sp and the significance of sclerite-covered taxa for early trochozoan evolution. SCIENTIFIC REPORTS[J]. 2017, 7(1): https://doaj.org/article/61eeb015bcd848c8b3e255f41bf775f4.
[46] 朱茂炎, 张俊明, 杨爱华, 李国祥, 赵方臣, 吕苗, 殷宗军. 华南新元古代地层、生－储－盖层发育与沉积环境. 中国东部中—新元古界地质学与油气资源. 2016, 107-135, http://ir.nigpas.ac.cn/handle/332004/12309.
[47] Yin, Zongjun, Zhu, Maoyan, Bottjer, David J, Zhao, Fangchen, Tafforeau, Paul. Meroblastic cleavage identifies some Ediacaran Doushantuo (China) embryo-like fossils as metazoans. GEOLOGY[J]. 2016, 44(9): 735-738, http://dx.doi.org/10.1130/G38262.1.
[48] Shixing Zhu, Maoyan Zhu, Andrew H Knoll, Zongjun Yin, Fangchen Zhao, Shufen Sun, Yuangao Qu, Min Shi, Huan Liu. Decimetre-scale multicellular eukaryotes from the 1.56-billion-year-old Gaoyuzhuang Formation in North China. NATURE COMMUNICATIONS[J]. 2016, 7(1): https://doaj.org/article/c24a1fdb24614cf9a52e5e788087010d.
[49] 朱茂炎, 张俊明, 杨爱华, 李国祥, 赵方臣, 吕苗, 殷宗军. 第5章：华南新元古代地层、生－储－盖层发育与沉积环境. 中国东部中-新元古界地质学与油气资源. 2016, 107-135, http://ir.nigpas.ac.cn/handle/332004/13076.
[50] Han Jian, Hu Shixue, Cartwright Paulyn, Zhao Fangchen, Ou Qiang, Kubota Shin, Wang Xing, Yang Xiaoguang. The earliest pelagic jellyfish with rhopalia from Cambrian Chengjiang Lagerstätte. PALAEOGEOGRAPHY, PALAEOCLIMATOLOGY, PALAEOECOLOGY[J]. 2016, 449: 166-173, http://dx.doi.org/10.1016/j.palaeo.2016.02.025.
[51] 杨爱华, 朱茂炎, 张俊明, 赵方臣, 吕苗. 扬子板块埃迪卡拉系（震旦系）陡山沱组层序地层划分与对比. 古地理学报[J]. 2015, 17(1): 1-1, http://ir.nigpas.ac.cn/handle/332004/11613.
[52] Ceng Han. First report of Wiwaxia from the Cambrian Chengjiang Lagerstätte. Geological Magazine. 2015, [53] 朱茂炎. A sponge grade body fossil with cellular resolution dating 60 Myr before the Cambrian. PNAS. 2015, [54] Zhao, Fangchen, Caron, JeanBernard, Bottjer, David J, Hu, Shixue, Yin, Zongjun, Zhu, Maoyan. Diversity and species abundance patterns of the Early Cambrian (Series 2, Stage 3) Chengjiang Biota from China. PALEOBIOLOGY[J]. 2014, 40(1): 50-69, https://www.webofscience.com/wos/woscc/full-record/WOS:000329529500005.
[55] Huang, Diying, Chen, Junyuan, Zhu, Maoyan, Zhao, Fangchen. The burrow dwelling behavior and locomotion of palaeoscolecidian worms: New fossil evidence from the Cambrian Chengjiang fauna. PALAEOGEOGRAPHY PALAEOCLIMATOLOGY PALAEOECOLOGY[J]. 2014, 398(398): 154-164, http://dx.doi.org/10.1016/j.palaeo.2013.11.004.
[56] Zeng, Han, Zhao, Fangchen, Yin, Zongjun, Li, Guoxiang, Zhu, Maoyan. A Chengjiang-type fossil assemblage from the Hongjingshao Formation (Cambrian Stage 3) at Chenggong, Kunming, Yunnan. CHINESE SCIENCE BULLETIN[J]. 2014, 59(25): 3169-3175, http://dx.doi.org/10.1007/s11434-014-0419-y.
[57] Zhao, Fangchen, Hu, Shixue, Zeng, Han, Zhu, Maoyan. A NEW HELMETIID ARTHROPOD FROM THE EARLY CAMBRIAN CHENGJIANG LAGERSTATTE, SOUTHWEST CHINA. JOURNAL OF PALEONTOLOGY[J]. 2014, 88(2): 367-370, https://www.webofscience.com/wos/woscc/full-record/WOS:000332752500015.
[58] 胡世学, 朱茂炎, 罗惠麟, 米歇尔.斯坦纳, 赵方臣, 李国祥, 刘琦, 张志飞. 关山生物群. 2013, 1-204, http://www.irgrid.ac.cn/handle/1471x/836925.
[59] Zhao, Fangchen, Bottjer, David J, Hu, Shixue, Yin, Zongjun, Zhu, Maoyan. Complexity and diversity of eyes in Early Cambrian ecosystems. SCIENTIFIC REPORTS[J]. 2013, 3(1): https://www.webofscience.com/wos/woscc/full-record/WOS:000324807000001.
[60] Zhu, Maoyan, Lu, Miao, Zhang, Junming, Zhao, Fangchen, Li, Guoxiang, Aihua, Yang, Zhao, Xin, Zhao, Meijuan. Carbon isotope chemostratigraphy and sedimentary facies evolution of the Ediacaran Doushantuo Formation in western Hubei, South China. PRECAMBRIAN RESEARCH[J]. 2013, 225(1): 7-28, http://dx.doi.org/10.1016/j.precamres.2011.07.019.
[61] Hu, Shixue, Steiner, Michael, Zhu, Maoyan, Luo, Huilin, Forchielli, Angela, Keupp, Helmut, Zhao, Fangchen, Liu, Qi. A new priapulid assemblage from the early Cambrian Guanshan fossil Lagerstatte of SW China. BULLETIN OF GEOSCIENCES[J]. 2012, 87(1): 93-106, http://ir.nigpas.ac.cn/handle/332004/4621.
[62] 赵方臣. A diverse and significant ichnofauna from the Lower Cambrian Wulongqing Formation near Kunming (Yunnan Province, South China). 2012, [63] Zhao, Fangchen, Hu, Shixue, Caron, JeanBernard, Zhu, Maoyan, Yin, Zongjun, Lu, Miao. Spatial variation in the diversity and composition of the Lower Cambrian (Series 2, Stage 3) Chengjiang Biota, Southwest China. PALAEOGEOGRAPHY PALAEOCLIMATOLOGY PALAEOECOLOGY[J]. 2012, 346: 54-65, http://dx.doi.org/10.1016/j.palaeo.2012.05.021.
[64] Lu, Miao, Zhu, Maoyan, Zhao, Fangchen. Revisiting the Tianjiayuanzi section - the stratotype section of the Ediacaran Doushantuo Formation, Yangtze Gorges, South China. BULLETIN OF GEOSCIENCES[J]. 2012, 87(1): 183-194, http://ir.nigpas.ac.cn/handle/332004/4624.
[65] 赵方臣. Revisit of the Tianjiayuanzi section: the stratotype section of Ediacaran Doushantuo Formation, Yangtze Gorges, South China. Bulletin of Geosciences. 2012, [66] 赵方臣, 朱茂炎, 胡世学. 不同生活方式的物种对快速沉积埋藏事件的反应——来自寒武纪早期澄江生物群中的证据. 古生物学报[J]. 2012, 51(3): 265-280, http://lib.cqvip.com/Qikan/Article/Detail?id=43324312.
[67] Zhu Maoyan, Yang Aihua, Zhao Fangchen, Zhang Junming, Zhao Yuanlong, Tai Tongshu. Cryogenian and cambrian successions in the slope facies of the Yangtze Platform at Wuhe, Jianhe County, Guizhou. 贵州大学学报 (自然科学版)[J]. 2012, 29(S1): 98-107, http://ir.nigpas.ac.cn/handle/332004/7967.
[68] 赵方臣. Widespread pyritization of insects from the Early Cretaceous Jehol biota. Palaios. 2012, [69] 王向东, 王玥, 张以春, 赵方臣, 盛青怡. 古生态学、古气候学、古生物地理学. 古生物学学科发展报告. 2010, 63-69, http://www.irgrid.ac.cn/handle/1471x/832892.
[70] Zhao FangChen, Zhu MaoYan, Hu ShiXue. Community structure and composition of the Cambrian Chengjiang biota. SCIENCE CHINA-EARTH SCIENCES[J]. 2010, 53(12): 1784-1799, http://www.irgrid.ac.cn/handle/1471x/474944.
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（1） 关山生物群——寒武纪早期海洋生物多样性的再现, The Guanshan Biota, 云南科技出版社, 2013-12, 第 5 作者

   

（ 1 ） 云南寒武纪澄江动物群埋藏学和定量古生态学研究, 主持, 国家级, 2011-01--2013-12
（ 2 ） 澄江动物群古群落分析, 主持, 省级, 2012-06--2015-06
（ 3 ） 澄江动物群区域埋藏相分析和古群落研究, 主持, 研究所（学校）, 2013-01--2015-12
（ 4 ） 从雪球事件到寒武纪大爆发--距今6 亿年前后的生物与环境演变, 参与, 国家级, 2013-01--2017-12
（ 5 ） 早期动物胚胎和成体化石同步辐射研究, 参与, 国家级, 2012-01--2014-12
（ 6 ） 多细胞生物起源与早期演化”子课题“瓮安生物群和澄江动物群特殊化石研, 参与, 部委级, 2011-01--2015-12
（ 7 ） 中科学院青年创新促进会, 主持, 部委级, 2014-01--2017-12
（ 8 ） 澄江生物群化石成因, 主持, 市地级, 2014-03--2016-03
（ 9 ） 澄江生物群、关山生物群和凯里生物群埋藏学及群落生态学对比研究, 主持, 国家级, 2015-01--2018-12
（ 10 ） 元古宙-显生宙转折期地球系统的剧烈波动与生物圈恢复力, 参与, 国家级, 2017-01--2020-12
（ 11 ） 新元古代-古生代转折期雪球事件和多细胞生物大爆发, 参与, 部委级, 2016-06--2020-06
（ 12 ） 早期生物群的演化, 参与, 部委级, 2017-05--2018-05
（ 13 ） 寒武纪特异埋藏化石群, 主持, 部委级, 2018-06--2023-06

（1）Complexity and diversity of eyes in Early Cambrian ecosystems: evidence from the Chengjiang Biota   Zhao F C   2013-09-21
（2）Diversity and species abundance patterns of the Early Cambrian Chengjiang Biota from China   Zhao F C   2013-09-21
（3） ‘Good vision’ triggered arthropods diversity and domination in the early Cambrian ecosystem   Zhao F C, Bottjer D J, Hu S X, Zhu M Y.   2013-04-06
（4）Comparative community paleoecology of the early Cambrian (Series 2, Stage 3) Chengjiang biota from China   ？ Zhao F C, Bottjer D J, Caron J B, Zhu M Y, Hu S X.   2012-11-03
（5）Community paleoecology of the early Cambrian Chengjiang Biota   Zhao F C、Zhu M Y、Hu S X   2012-06-09
（6）滇东地区寒武纪澄江动物群物种多样性空间变化特征   贵州关岭中国古生物学会第26届学术年会   赵方臣、胡世学、朱茂炎   2011-10-22

已指导学生

胡春林  硕士研究生  070903-古生物学与地层学  

曾晗  博士研究生  070903-古生物学与地层学